[1] 张煜星. 中国荒漠化气候类型的分布[J]. 干旱区研究, 1998, 15(2):46−50.

Zhang Y X. Distribution of climate type of Chinese desertification[J]. Arid Zone Research, 1998, 15(2): 46−50.
[2] 林年丰, 汤洁. 中国干旱半干旱区的环境演变与荒漠化的成因[J]. 地理科学, 2001, 21(1):24−29. doi:  10.3969/j.issn.1000-0690.2001.01.005

Lin N F, Tang J. Study on the environmental evolution and the causes of desertification in arid and semiarid regions in China[J]. Scientia Geographica Sinica, 2001, 21(1): 24−29. doi:  10.3969/j.issn.1000-0690.2001.01.005
[3] Barros V, Stocker T F. Managing the risks of extreme events and disasters to advance climate change adaptation: special report of the Intergovernmental Panel on Climate Change[J]. Journal of Clinical Endocrinology & Metabolism, 2012, 18(6): 586−599.
[4] Huang J, Yu H, Dai A, et al. Drylands face potential threat under 2 degrees C global warming target[J]. Nature Climate Change, 2017, 7(6): 417−422. doi:  10.1038/nclimate3275
[5] Li B, Chen Y, Shi X, et al. Temperature and precipitation changes in different environments in the arid region of northwest China[J]. Theoretical and Applied Climatology, 2013, 112(3−4): 589−596. doi:  10.1007/s00704-012-0753-4
[6] Huang J, Yu H, Guan X, et al. Accelerated dryland expansion under climate change[J]. Nature Climate Change, 2016, 6(2): 166−171. doi:  10.1038/nclimate2837
[7] Juan W Y, Cai R, Yun T, et al. Photosynthetic gas-exchange and PSⅡ photochemical acclimation to drought in a native and non-native xerophytic species (<italic>Artemisia ordosica</italic> and <italic>Salix psammophila</italic>)[J]. Ecological Indicators, 2018, 94: 130−138. doi:  10.1016/j.ecolind.2018.06.040
[8] 种培芳, 李毅, 苏世平. 荒漠植物红砂叶绿素荧光参数日变化及其与环境因子的关系[J]. 中国沙漠, 2010, 30(3):539−545.

Chong P F, Li Y, Su S P. Diurnal change in chlorophyllⅡfluorescence parameters of desert plant <italic>Reaumuria soongorica</italic> and its relationship with environmental factors[J]. Journal of Desert Research, 2010, 30(3): 539−545.
[9] 陈凤丽, 靳正忠, 李生宇, 等. 高温对花花柴(<italic>Karelinia caspica</italic>)光系统Ⅱ的影响[J]. 中国沙漠, 2013, 33(5):1371−1376. doi:  10.7522/j.issn.1000-694X.2013.00201

Chen F L, Jin Z Z, Li S Y, et al. Effects of heat stress on photosystem Ⅱ in <italic>Karelinia caspica</italic>[J]. Journal of Desert Research, 2013, 33(5): 1371−1376. doi:  10.7522/j.issn.1000-694X.2013.00201
[10] Chambers J C, Bradley B A, Brown C S, et al. Resilience to stress and disturbance, and resistance to <italic>Bromus tectorum</italic> L. invasion in cold desert shrublands of western North America[J]. Ecosystems, 2014, 17: 360−375. doi:  10.1007/s10021-013-9725-5
[11] Grant O M, Tronina L, García-Plazaola J L, et al. Resilience of a semi-deciduous shrub, <italic>Cistus salvifolius</italic>, to severe summer drought and heat stress[J]. Functional Plant Biology, 2014, 42(2): 219−228.
[12] 吴雅娟, 查天山, 贾昕, 等. 油蒿 (<italic>Artemisia ordosica</italic>) 光化学量子效率和非光化学淬灭的动态及其影响因子[J]. 生态学杂志, 2015, 34(2):319−325.

Wu Y J, Zha T S, Jia X, et al. Temporal variation and controlling factors of photochemical efficiency and non-photochemical quenching in <italic>Artemisia ordosica</italic>[J]. Chinese Journal of Ecology, 2015, 34(2): 319−325.
[13] 张继伟, 赵昕, 陈国雄, 等. 盐胁迫下荒漠植物柠条和油蒿的离子吸收及分配特征[J]. 干旱区资源与环境, 2016, 30(3):68−73.

Zhang J W, Zhao X, Chen G X, et al. Ion absorption and distribution in <italic>Caragana korshinskii</italic> and <italic>Artemisia ordosica</italic> seedlings under different NaCl stress[J]. Journal of Arid Land Resources and Environment, 2016, 30(3): 68−73.
[14] 王飞, 刘世增, 康才周, 等. 干旱胁迫对沙地云杉光合、叶绿素荧光特性的影响[J]. 干旱区资源与环境, 2017, 31(1):142−147.

Wang F, Liu S Z, Kang C Z, et al. Effects of drought stress on photosynthesis and chlorophyll fluorescence characteristics of <italic>Picea mongolica</italic>[J]. Journal of Arid Land Resources and Environment, 2017, 31(1): 142−147.
[15] 张明艳, 贾昕, 查天山, 等. 油蒿 (<italic>Artemisia ordosica</italic>) 光系统Ⅱ光化学效率对去除降雨的响应[J]. 中国沙漠, 2017, 37(3):475−482. doi:  10.7522/j.issn.1000-694X.2016.00021

Zhang M Y, Jia X, Zha T S, et al. PSⅡ photochemical efficiency of <italic>Artemisia ordosica</italic> in response to rainfall exclusion[J]. Journal of Desert Research, 2017, 37(3): 475−482. doi:  10.7522/j.issn.1000-694X.2016.00021
[16] 张景波, 张金鑫, 卢琦, 等. 乌兰布和沙漠油蒿叶片PSⅡ叶绿素荧光动力学参数及其光响应曲线动态[J]. 草业科学, 2019, 36(3):713−719.

Zhang J B, Zhang J X, Lu Q, et al. Dynamic changes of leaf parameters of PSⅡ fluorescence kinetics and fast photosynthetic response curves in <italic>Artemisia ordosica</italic>[J]. Pratacultural Science, 2019, 36(3): 713−719.
[17] 朴世龙, 张新平, 陈安平, 等. 极端气候事件对陆地生态系统碳循环的影响[J]. 中国科学:地球科学, 2019, 49(9):1321−1334. doi:  10.1360/N072018-00316

Piao S L, Zhang X P, Chen A P, et al. The impacts of climate extremes on the terrestrial carbon cycle: a review[J]. Scientia Sinica Terrae, 2019, 49(9): 1321−1334. doi:  10.1360/N072018-00316
[18] 任才, 贾昕, 吴雅娟, 等. 油蒿光系统Ⅱ光化学参数在生长季早期对降雪事件的响应[J]. 北京林业大学学报, 2019, 41(12):119−127. doi:  10.12171/j.1000-1522.20190058

Ren C, Jia X, Wu Y J, et al. Responses of PSⅡ photochemical parameter to a snowfall event in early growing season in <italic>Artemisia ordosica</italic>[J]. Journal of Beijing Forestry University, 2019, 41(12): 119−127. doi:  10.12171/j.1000-1522.20190058
[19] 杨洪晓, 张金屯, 吴波, 等. 油蒿对半干旱区沙地生境的适应及其生态作用[J]. 北京师范大学学报(自然科学版), 2004, 40(5):681−690.

Yang H X, Zhang J T, Wu B, et al. Adaptation of <italic>Artemisia ordosica</italic> to temperature arid sandy land and its roles in habitat shift[J]. Journal of Beijing Normal University (Natural Science), 2004, 40(5): 681−690.
[20] 曹生奎, 冯起, 司建华, 等. 植物水分利用效率研究方法综述[J]. 中国沙漠, 2009, 29(5):853−858.

Cao S K, Feng Q, Si J H, et al. Summary on research methods of water use efficiency in plant[J]. Journal of Desert Research, 2009, 29(5): 853−858.
[21] 王燕凌, 刘君, 李文兵, 等. 塔里木河下游刚毛柽柳光合作用、蒸腾作用及水分利用效率特性研究[J]. 新疆农业科学, 2015, 52(2):292−299.

Wang Y L, Liu J, Li W B, et al. Study on characteristics in photosynthesis, transpiration and water use efficiency of <italic>Tamarix hispida</italic> Willd. in the lower reaches of the Tarim River[J]. Xinjiang Agricultural Sciences, 2015, 52(2): 292−299.
[22] Massacci A, Nabiev S M, Pietrosanti L, et al. Response of the photosynthetic apparatus of cotton (<italic>Gossypium hirsutum</italic>) to the onset of drought stress under field conditions studied by gas-exchange analysis and chlorophyll fluorescence imaging[J]. Plant Physiology and Biochemistry, 2008, 46(2): 189−195. doi:  10.1016/j.plaphy.2007.10.006
[23] 张亚娟, 谢忠奎, 赵学勇, 等. 水分胁迫对东方百合光合特性、叶绿素荧光参数及干物质积累的影响[J]. 中国沙漠, 2011, 31(4):884−888.

Zhang Y J, Xie Z K, Zhao X Y, et al. Effects of water stress on photosynthetic characteristics, chlorophyll fluorescence, and dry matter of <italic>Oriental lilies</italic>[J]. Journal of Desert Research, 2011, 31(4): 884−888.
[24] Xu W Z, Deng X P, Xu B C. Photosynthetic activity and efficiency of <italic>Bothriochloa ischaemum</italic> and <italic>Lespedeza davurica</italic> in mixtures across growth periods under water stress[J]. Acta Physiologiae Plantarum, 2014, 36(4): 1033−1044. doi:  10.1007/s11738-013-1481-9
[25] 贾虎森, 韩亚琴. 高等植物光合作用的光抑制研究进展[J]. 植物学通报, 2000, 17(3):218−224.

Jia H S, Han Y Q. Advances in studies on photoinhibition in photosynthesis of higher plants[J]. Chinese Bulletin of Botany, 2000, 17(3): 218−224.
[26] 石莎, 冯金朝, 邹学勇. 腾格里沙漠南缘2种沙地灌木植物的光合特征[J]. 云南大学学报(自然科学版), 2007, 29(5):519−524. doi:  10.3321/j.issn:0258-7971.2007.05.019

Shi S, Feng J Z, Zou X Y. Photosynthetic characteristics of two desert shrubs in south of Tengger Desert[J]. Journal of Yunnan University (Natural Sciences Edition), 2007, 29(5): 519−524. doi:  10.3321/j.issn:0258-7971.2007.05.019
[27] 韩刚, 赵忠. 不同土壤水分下4种沙生灌木的光合光响应特性[J]. 生态学报, 2010, 30(15):4019−4026.

Han G, Zhao Z. Light response characteristics of photosynthesis of four xerophilous shrubs under different soil moistures[J]. Acta Ecologica Sinica, 2010, 30(15): 4019−4026.
[28] Doupis G, Bertaki M, Psarras G, et al. Water relations, physiological behavior and antioxidant defence mechanism of olive plants subjected to different irrigation regimes[J]. Scientia Horticulturae, 2013, 153(3): 150−156.
[29] 何炎红, 白玉娥, 王海燕, 等. 光胁迫对沙冬青叶绿素荧光特征和光呼吸的影响[J]. 西北农业学报, 2015, 24(10):124−130. doi:  10.7606/j.issn.1004-1389.2015.10.018

He Y H, Bai Y E, Wang H Y, et al. Effect of light stress on chlorophyll fluorescence and photorespiration of <italic>Ammopiptanthus mongolicus</italic>[J]. Acta Agriculturae Boreali-Occidentalis Sinica, 2015, 24(10): 124−130. doi:  10.7606/j.issn.1004-1389.2015.10.018
[30] 姜闯道, 高辉远, 邹琦, 等. 叶角、光呼吸和热耗散协同作用减轻大豆幼叶光抑制[J]. 生态学报, 2005, 25(2):319−325. doi:  10.3321/j.issn:1000-0933.2005.02.021

Jiang C D, Gao H Y, Zou Q, et al. The co-operation of leaf orientation, photorespiration and thermal dissipation alleviate photoinhibition in young leaves of soybean plants[J]. Acta Ecologica Sinica, 2005, 25(2): 319−325. doi:  10.3321/j.issn:1000-0933.2005.02.021
[31] Öquist G, Chow W S, Anderson J M. Photoinhibition of photosynthesis represents a mechanism for the long-term regulation of photosystem Ⅱ[J]. Planta, 1992, 186(3): 450−460.
[32] Ruban A V, Murchie E H. Assessing the photoprotective effectiveness of non-photochemical chlorophyll fluorescence quenching: a new approach[J]. Biochimica et Biophysica Acta (BBA)-Bioenergetics, 2012, 1817(7): 977−982. doi:  10.1016/j.bbabio.2012.03.026
[33] Ruban A V. Nonphotochemical chlorophyll fluorescence quenching: mechanism and effectiveness in protecting plants from photodamage[J]. Plant Physiology, 2016, 170(4): 1903−1916. doi:  10.1104/pp.15.01935
[34] 阮成江, 李代琼. 黄土丘陵区沙棘气孔导度及其影响因子[J]. 西北植物学报, 2001, 21(6):30−36.

Ruan C J, Li D Q. Stomatal conductance and influence factors of seabuckthorn in loess hilly region[J]. Acta Botanica Boreali-Occidentalia Sinica, 2001, 21(6): 30−36.
[35] Campos H, Trejo C, Peña-Valdivia C B, et al. Stomatal and non-stomatal limitations of bell pepper (<italic>Capsicum annuum</italic> L <italic></italic>.) plants under water stress and re-watering: delayed restoration of photosynthesis during recovery[J]. Environmental & Experimental Botany, 2014, 98(1): 56−64.
[36] 许大全. 光系统Ⅱ反应中心的可逆失活及其生理意义[J]. 植物生理学通讯, 1999, 35(4):273−276.

Xu D Q. Reversible inactivation of photosystem Ⅱ reaction centers and its physiological ignificance[J]. Plant Physiology Communications, 1999, 35(4): 273−276.
[37] Porcar-Castell A, Pfündel E, Korhonen J F J, et al. A new monitoring PAM fluorometer (MONI-PAM) to study the short-and long-term acclimation of photosystem Ⅱ in field conditions[J]. Photosynthesis Research, 2008, 96(2): 173−179. doi:  10.1007/s11120-008-9292-3
[38] Kalaji H M, Jajoo A, Oukarroum A, et al. Chlorophyll a fluorescence as a tool to monitor physiological status of plants under abiotic stress conditions[J]. Acta Physiologiae Plantarum, 2016, 38: 102−112. doi:  10.1007/s11738-016-2113-y
[39] 靳虎甲, 王继和, 李毅, 等. 油蒿生态学研究综述[J]. 西北林学院学报, 2009, 24(4):62−66.

Jin H J, Wang J H, Li Y, et al. Summary of <italic>Artemisia ordosica</italic> ecology studies[J]. Journal of Northwest Forestry University, 2009, 24(4): 62−66.
[40] 马全林, 郑庆中, 贾举杰, 等. 乌兰布和沙漠沙蒿与油蒿群落的物种组成与数量特征[J]. 生态学报, 2012, 3(11):3423−3431.

Ma Q L, Zheng Q Z, Jia J J, et al. Quantitative characteristics and species composition of <italic>Artemisia sphaerocephala</italic> and <italic>A. ordosica</italic> communities in the Ulanbuh Desert[J]. Acta Ecologica Sinica, 2012, 3(11): 3423−3431.
[41] Jia X, Zha T S, Wu B, et al. Biophysical controls on net ecosystem CO<sub>2</sub> exchange over a semiarid shrubland in northwest China[J]. Biogeosciences Discussions, 2014, 11(3): 4679−4693.
[42] Schansker G, Van Rensen J J S. Performance of active photosystem Ⅱ centers in photoinhibited pea leaves[J]. Photosynthesis Research, 1999, 62(2): 175−184.
[43] 刘晓晴, 常宗强, 马亚丽, 等. 胡杨(<italic>Populus euphratica</italic>)异形叶叶绿素荧光动力学[J]. 中国沙漠, 2014, 34(3):704−711. doi:  10.7522/j.issn.1000-694X.2014.00007

Liu X Q, Chang Z Q, Ma Y L, et al. Characteristics of the fast chlorophyll fluorescence induction kinetics of heteromorphic leaves in <italic>Populus euphratica</italic>[J]. Journal of Desert Research, 2014, 34(3): 704−711. doi:  10.7522/j.issn.1000-694X.2014.00007
[44] 师生波, 张怀刚, 师瑞, 等. 青藏高原春小麦叶片光合作用的光抑制及PSⅡ反应中心光化学效率的恢复分析[J]. 植物生态学报, 2014, 38(4):375−386. doi:  10.3724/SP.J.1258.2014.00034

Shi S B, Zhang H G, Shi R, et al. Assessment of photosynthetic photo-inhibition and recovery of PSⅡ photochemical efficiency in leaves of wheat varieties in Qinghai-Xizang Plateau[J]. Chinese Journal of Plant Ecology, 2014, 38(4): 375−386. doi:  10.3724/SP.J.1258.2014.00034
[45] 郝建卿, 吕娜, 杨扬, 等. 内蒙乌海胡杨异形叶水分及叶绿素荧光参数的比较[J]. 北京林业大学学报, 2010, 32(5):41−44.

Hao J Q, Lü N, Yang Y, et al. Comparative study of chlorophyll fluorescence parameters and water physiological characters of heteromorphic leaves for <italic>Populus euphratica</italic>[J]. Journal of Beijing Forestry University, 2010, 32(5): 41−44.
[46] 韩婧, 吴益, 赵琳,等. 光周期对促成栽培芍药生长开花和叶绿素荧光动力学影响[J]. 北京林业大学学报, 2015, 37(9):62−69.

Han J, Wu Y, Zhao L, et al. Effects of photoperiod on the growth, flowering and chlorophyll fluorescence kinetics of forced <italic>Paeonia lactiflora</italic>[J]. Journal of Beijing Forestry University, 2015, 37(9): 62−69.