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广东南岭国家级自然保护区华南五针松种内和种间关系研究

王俊 王泳腾 段洋波 谢磊 崔国发

王俊, 王泳腾, 段洋波, 谢磊, 崔国发. 广东南岭国家级自然保护区华南五针松种内和种间关系研究[J]. 北京林业大学学报, 2020, 42(5): 25-32. doi: 10.12171/j.1000-1522.20180436
引用本文: 王俊, 王泳腾, 段洋波, 谢磊, 崔国发. 广东南岭国家级自然保护区华南五针松种内和种间关系研究[J]. 北京林业大学学报, 2020, 42(5): 25-32. doi: 10.12171/j.1000-1522.20180436
Wang Jun, Wang Yongteng, Duan Yangbo, Xie Lei, Cui Guofa. Interspecific and intraspecific competitions of Pinus kwangtungensis in Nanling National Nature Reserve, Guangdong Province of southern China[J]. Journal of Beijing Forestry University, 2020, 42(5): 25-32. doi: 10.12171/j.1000-1522.20180436
Citation: Wang Jun, Wang Yongteng, Duan Yangbo, Xie Lei, Cui Guofa. Interspecific and intraspecific competitions of Pinus kwangtungensis in Nanling National Nature Reserve, Guangdong Province of southern China[J]. Journal of Beijing Forestry University, 2020, 42(5): 25-32. doi: 10.12171/j.1000-1522.20180436

广东南岭国家级自然保护区华南五针松种内和种间关系研究

doi: 10.12171/j.1000-1522.20180436
基金项目: 北京林业大学青年教师中长期科研项目(2015ZCQ-BH-03),黄淮学院青年骨干教师资助计划,黄淮学院合格课程(1501HK168)
详细信息
    作者简介:

    王俊。主要研究方向:生物多样性保育。Email:281879789@qq.com  地址:100083北京市海淀区北京林业大学生态与自然保护学院

    责任作者:

    崔国发,教授,博士生导师。主要研究方向:自然保护区学。Email:fa6716@163.com  地址:同上

  • 中图分类号: S718.5

Interspecific and intraspecific competitions of Pinus kwangtungensis in Nanling National Nature Reserve, Guangdong Province of southern China

  • 摘要: 目的  通过对广东南岭国家级自然保护区小黄山内118株对象木及253株伴生种的调查,分析华南五针松的竞争强度,揭示华南五针松群落的动态变化规律和种群生态适应机制。 方法  沿海拔梯度设置标准群落样方。利用Hegyi单木竞争模型计算华南五针松群落内物种间的竞争指数,并采用回归分析的方法建立对象木胸径与林分、种间竞争强度的函数模型。 结果  (1)华南五针松胸径与树高存在显著的幂函数关系;(2)华南五针松的种内和种间竞争指数分别为893.736和559.682;(3)华南五针松伴生种较多,其主要种间竞争树种的顺序为木荷 > 甜槠 > 五列木 > 马尾松 > 长苞铁杉 > 凤凰木 > 福建柏 > 马蹄荷;(4)华南五针松胸径与整个林分和种间竞争指数呈显著的负相关。 结论  华南五针松群落内以种内竞争为主。竞争强度随华南五针松个体胸径的增大而减小,当华南五针松胸径到达30 cm时,竞争强度下降较为平稳,二者符合幂函数关系,所得模型能有效的预测华南五针松的种内和种间强度。

     

  • 图  1  研究区华南五针松胸径与树高的关系

    Figure  1.  Relationship between DBH and tree height of Pinus kwangtungensis in study area

    图  2  华南五针松胸径与整个林分的竞争指数关系

    Figure  2.  Relationship between DBH of P. kwangtungensis and CI of the overall stand

    图  3  华南五针松胸径与伴生树种的竞争指数关系

    Figure  3.  Relationship between DBH of P. kwangtungensis and CI of associated tree species

    表  1  调查样地的基本概况

    Table  1.   General description of the sample plots

    样地编号
    Sample plot No.
    地点
    Site
    经纬度
    Longitude and latitude
    海拔
    Altitude/m
    坡度
    Gradient/(°)
    坡向
    Slope aspect
    郁闭度
    Canopy density
    林分起源
    Stand origin
    YD001 乳峰楼
    Rufenglou
    24°53′56″N,113°01′11″E 1 575 60 ~ 65 正南
    Due south
    0.80 天然林
    Natural forest
    YD002 乳阳小黄山
    Ruyang Xiaohuang Mountain
    24°53′52″N,113°01′13″E 1 509 37 ~ 43 正南
    Due south
    0.82 天然林
    Natural forest
    YD003 乳阳小黄山
    Ruyang Xiaohuang Mountain
    24°53′58″N,113°01′22″E 1 447 50 ~ 60 正南
    Due south
    0.83 天然林
    Natural forest
    YD004 乳阳小黄山
    Ruyang Xiaohuang Mountain
    24°53′45″N,113°01′40″E 1 186 50 ~ 55 东南
    Southeast
    0.78 天然林
    Natural forest
    YD005 乳阳小黄山
    Ruyang Xiaohuang Mountain
    24°53′49″N,113°01′34″E 1 276 60 ~ 65 东南
    Southeast
    0.75 天然林
    Natural forest
    YD006 乳阳小黄山
    Ruyang Xiaohuang Mountain
    24°53′51″N,113°01′29″E 1 370 45 ~ 50 东北
    Northeast
    0.80 天然林
    Natural forest
    YD007 瀑布亭
    Pubuting
    24°53′49″N,113°02′29″E 994 51 ~ 56 东南
    Southeast
    0.60 天然林
    Natural forest
    下载: 导出CSV

    表  2  对象木与竞争木概况

    Table  2.   Status of obiective trees and competitive trees

    径级
    DBH class/cm
    对象木 Objective tree竞争木 Competitive tree
    株数
    Plant number
    平均胸径
    Average DBH/cm
    平均树高
    Average tree height/m
    株数
    Plant number
    平均胸径
    Average DBH/cm
    平均树高
    Average tree height/m
    < 5 0 0.000 0.000 4 4.400 5.300
    5 ~ 10 22 8.104 7.700 49 8.000 7.537
    10 ~ 15 24 12.346 9.680 72 12.744 10.071
    15 ~ 20 20 17.680 11.310 47 17.860 10.991
    20 ~ 25 7 23.270 17.260 24 22.754 13.321
    25 ~ 30 6 27.900 15.317 19 27.774 14.611
    30 ~ 35 12 32.800 19.917 17 32.341 17.400
    35 ~ 40 10 37.550 20.850 14 37.950 17.743
    > 40 17 58.047 21.306 7 52.771 15.200
    合计 Total 118 253
    下载: 导出CSV

    表  3  华南五针松种内和种间竞争强度

    Table  3.   Intraspecific and interspecific competition of P. kwangtungensis

    径级
    DBH class/cm
    株数
    Plant number
    种内竞争
    Intraspecific competition
    种间竞争
    Interspecific competition
    竞争指数
    Competitive index (CI)
    平均竞争指数
    Average CI
    竞争指数
    CI
    平均竞争指数
    Average CI
    < 5 0 0.000 0.000 0.000 0.000
    5 ~ 10 22 425.009 19.319 238.994 10.818
    10 ~ 15 24 253.314 10.555 155.335 6.472
    15 ~ 20 20 103.273 3.740 74.803 8.904
    20 ~ 25 7 37.416 5.345 22.474 3.211
    25 ~ 30 6 14.353 2.392 12.962 2.160
    30 ~ 35 12 30.050 2.504 21.689 1.807
    35 ~ 40 10 23.415 2.342 15.964 1.596
    > 40 17 6.906 0.406 17.461 1.027
    合计 Total 118 893.736 559.682
    下载: 导出CSV

    表  4  华南五针松竞争木的种类组成和竞争强度

    Table  4.   Species composition of competitive trees and competition intensities for Pinus kwangtungensis

    树种
    Tree species
    株数
    Plant number
    比例
    Percentage/%
    平均胸径
    Average DBH/cm
    平均树高
    Average tree height/m
    竞争指数
    CI
    平均竞争指数
    Average CI
    排序
    Ranking
    木荷 Schima superba 39 15.42 17.713 12.382 210.592 5.400 1
    甜槠 Castanopsis eyrei 40 15.80 29.103 12.815 107.057 2.676 2
    五列木 Pentaphylax euryoidex 42 16.60 12.843 9.936 61.107 1.455 3
    马尾松 Pinus massoniana 6 2.37 23.717 17.417 33.247 5.541 4
    长苞铁杉 Tsuga longibracteata 11 4.35 23.091 13.827 27.991 2.545 5
    凤凰木 Delonix regia 2 0.79 9.550 10.000 20.039 10.020 6
    福建柏 Fokienia hodginsii 16 6.32 17.938 9.744 18.292 1.143 7
    马蹄荷 Exbucklandia populnea 9 3.56 14.444 10.778 15.813 1.757 8
    石壁杜鹃 Rhododendron bachii 4 1.58 11.600 5.875 11.218 2.805 9
    厚叶冬青 Ilex elmerrilliana 2 0.79 6.450 5.600 9.589 4.795 10
    红淡比 Cleyera japonica 1 0.40 4.900 6.700 7.132 7.132 11
    青冈 Cyclobalanopsis glauca 12 4.74 19.558 12.542 7.124 0.594 12
    南方铁杉 Tsuga chinensis 5 1.98 17.160 7.620 6.876 1.375 13
    金叶含笑 Michelia foveolata 5 1.98 17.660 8.420 4.658 0.932 14
    黄背青冈 Cyclobalanopsis poilanei 4 1.58 20.950 10.000 3.435 0.859 15
    乐东拟单性木兰 Parakmeria lotungensis 4 1.58 29.575 18.575 2.382 0.596 16
    羊角杜鹃 Rhododendron cavaleriei 9 3.56 10.856 7.544 1.748 0.194 17
    新木姜子 Neolitsea aurata 1 0.40 8.500 6.500 1.547 1.547 18
    杨桐 Adinandra millettii 4 1.58 20.625 11.750 1.483 0.371 19
    杨梅 Myrica rubra 2 0.79 24.400 14.000 1.424 0.712 20
    凤凰木 Delonix regia 2 0.79 11.150 9.150 1.078 0.539 21
    其他26种 Other twenty-six species 33 13.04 14.807 10.616 5.851 0.177
    合计 Total 253 1 559.682
    下载: 导出CSV

    表  5  华南五针松胸径与竞争强度的模型参数

    Table  5.   Model parameters of competition intensity and DBH of P. kwangtungensis

    项目 Item类别 Sort显著性
    Significance
    ABR2
    华南五针松与伴生树种 P. kwangtungensis and associated tree species 138.22 1.292 0.672 P < 0.01
    华南五针松与整个林分 P. kwangtungensis and overall stand 668.34 1.547 0.674 P < 0.01
    注:AB为回归模型(幂函数)CI = AD− B的参数。Notes: A and B are the parameters of regression model (power function) CI = AD− B.
    下载: 导出CSV

    表  6  华南五针松竞争强度与个体胸径的模型预测

    Table  6.   Model prediction of individual DBH and competitive intensity of P. kwangtungensis

    项目 Item径级 DBH/cm
    0 ~ 55 ~ 1010 ~ 1515 ~ 2020 ~ 2525 ~ 3030 ~ 3535 ~ 40> 40
    华南五针松与整个林分 P. kwangtungensis and overall stand 161.95 26.26 13.69 7.85 5.14 3.88 3.02 2.45 1.25
    华南五针松与伴生树种 P. kwangtungensis and associated tree species 20.38 9.41 5.16 3.34 2.44 1.89 1.55 1.26 0.82
    下载: 导出CSV
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出版历程
  • 收稿日期:  2019-01-03
  • 修回日期:  2019-05-21
  • 网络出版日期:  2020-05-11
  • 刊出日期:  2020-07-01

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