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吉林蛟河针阔混交林幼苗动态及环境驱动因子

李冠男 张春雨 赵秀海

李冠男, 张春雨, 赵秀海. 吉林蛟河针阔混交林幼苗动态及环境驱动因子[J]. 北京林业大学学报, 2021, 43(8): 41-49. doi: 10.12171/j.1000-1522.20200356
引用本文: 李冠男, 张春雨, 赵秀海. 吉林蛟河针阔混交林幼苗动态及环境驱动因子[J]. 北京林业大学学报, 2021, 43(8): 41-49. doi: 10.12171/j.1000-1522.20200356
Li Guannan, Zhang Chunyu, Zhao Xiuhai. Seedling dynamics and environmental driving factors of coniferous and broadleaved mixed forest in Jiaohe, Jilin Province of northeastern China[J]. Journal of Beijing Forestry University, 2021, 43(8): 41-49. doi: 10.12171/j.1000-1522.20200356
Citation: Li Guannan, Zhang Chunyu, Zhao Xiuhai. Seedling dynamics and environmental driving factors of coniferous and broadleaved mixed forest in Jiaohe, Jilin Province of northeastern China[J]. Journal of Beijing Forestry University, 2021, 43(8): 41-49. doi: 10.12171/j.1000-1522.20200356

吉林蛟河针阔混交林幼苗动态及环境驱动因子

doi: 10.12171/j.1000-1522.20200356
基金项目: 国家自然科学基金项目(31971650),国家重点研发计划重点专项(2017YFC0504104)
详细信息
    作者简介:

    李冠男。主要研究方向:森林经营理论与技术。Email:guannan_li@outlook.com 地址:100083 北京市海淀区清华东路35号北京林业大学林学院

    责任作者:

    张春雨,博士,教授。主要研究方向:森林经营理论与技术、生物多样性与生态系统功能。Email:zcy_0520@163.com 地址:同上

  • 中图分类号: S754.1

Seedling dynamics and environmental driving factors of coniferous and broadleaved mixed forest in Jiaohe, Jilin Province of northeastern China

  • 摘要:   目的  本文以吉林蛟河不同发育阶段针阔混交林为研究对象,对比分析幼苗密度的核密度估计曲线年际变化规律,探讨土壤因子的边际变化和土壤因子的分布变化对幼苗密度分布动态的相对影响。  方法  在420 m × 520 m的中龄林样地和500 m × 840 m的成熟林样地中系统布设幼苗调查样方。利用分位数回归和反事实分解法检验环境因子在不同分位数水平上对幼苗密度的边际影响,进而明确幼苗密度在不同分位点上呈现不均衡变化的主导因素。  结果  幼苗密度的核密度估计曲线呈峰值向左偏移、长尾向右侧延伸的正偏态分布。在θ = 0.90的高分位点,中龄林样地中幼苗密度变化值为−5.9,成熟林样地幼苗密度变化值为−2.6。中龄林样地在θ = 0.75时幼苗密度变化值为5.8,成熟林样地在θ = 0.50和θ = 0.75时幼苗密度变化值均为2。在高分位点上幼苗密度的更大变化反映了右单尾概率分布的不均衡性。在不同估计分位点上系数效应和协变量效应对幼苗密度分布变化的相对作用大小不同。中龄林样地中系数效应在全部估计分位点上都具有很高的解释量;成熟林样地中在θ = 0.50分位点上,协变量效应对幼苗密度变化具有89%的解释量,在其他的估计分位点上系数效应的解释量更高。因此,土壤因子分布变化对幼苗密度分布变化影响的相对作用更大,而土壤因子边际变化则是导致幼苗密度不均衡变化的主要因素。土壤中速效氮、速效磷、速效钾分布变化对每个估计分位点上幼苗密度变化的解释量大多不足30%,而土壤含水量和土壤pH值对幼苗密度变化具有相对更大的影响。  结论  幼苗密度的年际变化在不同分位点上是不均衡的,在高分位点上的变化尤为明显。土壤因子的边际变化和土壤因子的分布变化共同决定了幼苗的存活动态。

     

  • 图  1  样地中幼苗监测点的设置示意图

    Figure  1.  Layout of monitoring stations in forest sample plots

    图  2  幼苗密度的核密度估计值的年际变化

    A ~ C为中龄林林地,D ~ F为成熟林林地。A−C represent HF sample plot, D−F represent MF sample plot.

    Figure  2.  Comparison of kernel density estimates of number distribution between 2012 and 2013

    图  3  中龄林样地中幼苗密度动态与环境变量的分位数回归

    Figure  3.  Quantile regression between seedling dynamics and environmental variables in HF sample plots

    图  4  成熟林样地中幼苗密度动态与环境变量的分位数回归

    Figure  4.  Quantile regression between seedling dynamics and environmental variables in MF sample plots

    表  1  中龄林幼苗的物种组成动态

    Table  1.   Quantitative composition of seedling species in half-matured forest (HF)

    树种
    Tree species
    2012年苗数
    Seedling number
    in 2012
    2013年苗数
    Seedling number
    in 2013
    2012—2013年
    新生苗数
    Recruitment number
    in 2012−2013
    2012—2013年
    死亡苗数
    Dead seedling number
    in 2012−2013
    2012—2013年
    死亡率
    Mortality rate in
    2012−2013/%
    2012年
    重要值
    Importance value
    (IV) in 2012
    2013年
    重要值
    IV in 2013
    水曲柳
    Fraxinus mandschurica
    677 311 218 584 86.3 52.96 31.00
    沙松
    Abies holophylla
    311 45 10 276 88.7 17.87 4.72
    色木槭
    Acer mono
    118 446 409 81 68.6 13.78 38.22
    白牛槭
    Acer mandshuricum
    55 369 353 39 70.9 7.87 33.43
    胡桃楸
    Juglans mandshurica
    46 7 7 46 100.0 5.26 1.02
    红松
    Pinus koraiensis
    28 5 5 28 100.0 2.19 0.57
    春榆
    Ulmus davidiana var.
    japonica
    27 32 9 4 14.8 2.28 3.60
    簇毛槭
    Acer barbinerve
    23 10 10 23 100.0 1.75 1.13
    青杨
    Populus cathayana
    23 36 22 9 39.1 1.24 1.60
    紫椴
    Tilia amurensis
    10 39 39 10 100.0 1.01 3.61
    裂叶榆
    Ulmus laciniata
    4 2 0 2 50.0 0.53 0.33
    黄檗
    Phellodendron amurense
    3 0 0 3 100.0 0.37 0.00
    青楷槭
    Acer tegmentosum
    3 23 23 3 100.0 0.37 2.51
    稠李
    Padus racemosa
    1 0 0 1 100.0 0.16 0.00
    山杨
    Populus davidiana
    1 0 0 1 100.0 0.16 0.00
    鼠李
    Rhamnus davurica
    1 0 0 1 100.0 0.16 0.00
    糠椴
    Tilia mandshurica
    0 1 1 0 0.0 0.00 0.16
    千金榆
    Carpinus cordata
    0 5 5 0 0.0 0.00 0.82
    合计 Total 1 331 1 331 1 111 1 111 83.5
    下载: 导出CSV

    表  2  成熟林幼苗的物种组成动态

    Table  2.   Quantitative composition of seedling species in mature forest (MF)

    树种
    Tree species
    2012年苗数
    Seedling number
    in 2012
    2013年苗数
    Seedling number
    in 2013
    2012—2013年
    新生苗
    Recruitment number
    in 2012−2013
    2012—2013年
    死亡苗数
    Dead seedling number
    in 2012−2013
    2012—2013年
    死亡率
    Mortality rate in
    2012−2013/%
    2012年
    重要值
    IV in 2012
    2013年
    重要值
    IV in 2013
    水曲柳
    Fraxinus mandschurica
    1403 537 219 1085 77.3 40.39 24.32
    沙松
    Abies holophylla
    1391 41 3 1353 97.3 29.25 2.11
    色木槭
    Acer mono
    538 1481 1402 459 85.3 22.40 50.23
    白牛槭
    Acer mandshuricum
    143 750 724 117 81.8 8.42 30.06
    糠椴
    Tilia mandshurica
    118 104 104 118 100.0 5.38 5.23
    红松
    Pinus koraiensis
    89 30 27 86 96.6 4.38 1.54
    暴马丁香
    Syringa reticulata
    264 0 0 264 100.0 3.96 0.00
    紫椴
    Tilia amurensis
    80 435 420 65 81.3 3.80 14.94
    簇毛槭
    Acer barbinerve
    131 45 30 116 88.5 3.73 2.28
    裂叶榆
    Ulmus laciniata
    29 2 0 27 93.1 1.71 0.15
    胡桃楸
    Juglans mandshurica
    27 8 5 24 88.9 1.69 0.59
    千金榆
    Carpinus cordata
    26 87 84 23 88.5 1.56 5.12
    春榆
    Ulmus davidiana var.
    japonica
    24 47 27 4 16.7 1.36 2.78
    青楷槭
    Acer tegmentosum
    23 130 128 21 91.3 1.10 7.13
    蒙古栎
    Quercus mongolica
    10 3 2 9 90.0 0.71 0.22
    柠筋槭
    Acer triflorum
    6 20 15 1 16.7 0.43 1.35
    毛榛
    Corylus mandshurica
    3 0 0 3 100.0 0.09 0.00
    黄檗
    Phellodendron amurense
    1 0 0 1 100.0 0.07 0.00
    合计 Total 4 306 3 720 3 190 3 776 87.7
    下载: 导出CSV

    表  3  不同分位点上幼苗密度分布变化的反事实分解

    Table  3.   Decomposing the changes in kernel density of number at each estimated quantiles

    样地
    Sample plot
    分位点
    Quantile (θ)
    边际分布
    Marginal distribution
    解释比例
    Explained proportion
    单个协变量对变化量的解释比例
    Proportion of overall changes explained by an individual covariate
    20122013变化量
    Change
    协变量效应
    Covariate
    effect
    系数效应
    Coefficient
    effect
    土壤含水量
    Soil moisture
    content
    土壤酸碱度
    Soil pH
    碱解氮
    Alkali hydrolyzed
    nitrogen
    速效磷
    Available
    phosphorus
    速效钾
    Available
    potassium
    中龄林样地
    HF sample plot
    0.104.04.70.70.100.900.060.030.010.25
    0.257.37.40.10.040.960.670.440.25
    0.5011.515.03.50.030.970.300.02
    0.7518.023.85.80.130.860.100.08
    0.9037.231.3−5.91.000.240.28
    成熟林样地
    MF sample plot
    0.103.04.01.00.040.960.330.730.300.04
    0.257.07.20.20.070.930.340.980.570.490.04
    0.5012.014.02.00.890.110.31
    0.7522.024.02.00.040.960.270.480.980.680.07
    0.9042.239.6−2.60.470.530.56
    下载: 导出CSV
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出版历程
  • 收稿日期:  2020-11-17
  • 修回日期:  2021-01-02
  • 网络出版日期:  2021-06-19
  • 刊出日期:  2021-08-31

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