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    黄土高原地区4种常见树种适宜区对气候变化响应

    Response of four common tree species suitable areas to climate change in the Loess Plateau region of northern China

    • 摘要:
        目的  分析黄土高原地区4个常见树种(樟子松、油松、柠条、华北落叶松)在当前和未来的潜在分布,揭示气候变化对植物空间分布格局的影响。
        方法  基于19个气候因子结合来自Ho1dridge生命地带模型和Kira指标体系的5个指标:年生物温度(ABT)、潜在蒸散率(PER)、温暖指数(WI)、寒冷指数(CI)、干燥指数(HI),运用Maxent模型,模拟预测了4个树种在当前、未来(2041—2060年、2061—2080年)ssp126、ssp245、ssp585气候情景下的潜在地理分布。运用刀切图分析影响其分布的主要环境因子,并采用受试者工作特征曲线(ROC)下的面积(AUC)对预测结果进行检验。
        结果  (1)Maxent模型可以较好地模拟黄土高原4个主要常见种的地理分布范围,各物种的10次平均AUC结果均大于0.8;(2)对于樟子松、油松、柠条来说,温度和水分共同限制其分布,而对于华北落叶松来说,影响其分布的主导因子是降水。温度季节性变化标准差、潜在蒸散率、最湿润季节降雨量、最干燥月降水都影响樟子松分布。年均温变化范围、温度季节性变化标准差、最冷月最低温度,最暖季度降水量、潜在蒸散率是影响油松分布的主导因子。影响柠条适宜区分布的主导因子为最暖月最高温度、温暖指数、等温性和潜在蒸散率、最冷季度降水量。影响华北落叶松分布的主导因子主要与降水有关,分别是最湿润月降雨量、最湿润季节降雨量、降水量变异系数和潜在蒸散率。(3)油松、柠条和华北落叶松的潜在适生区将向西北方向迁移,樟子松的潜在适生区向西南方向迁移。油松和华北落叶松的潜在适宜区面积呈现先扩大后缩小的趋势,而柠条和樟子松的潜在适生区将持续扩张,尤其是樟子松的高适生区占比将在2070s扩大至50.97%。
        结论  气候变化将使油松和华北落叶松丧失一部分的高适生区,但同时会使柠条和樟子松的高适生区扩张明显。在黄土高原退耕还林建设中,可优先考虑种植柠条和樟子松。

       

      Abstract:
        Objective  This paper aims to analyze the current and future potential distributions of four common tree species (Pinus sylvestris, Pinus tabuliformis, Caragana korshinskii and Larix gmelinii var. principis-rupprechtii) in the Loess Plateau region of northern China, and to reveal the effects of climate change on spatial distribution patterns of plants.
        Method  Four dominant species on the Loess Plateau were studied based on 19 climate factors and 5 indicators from Holdridge life zone model and Kira index system: bio-temperature (ABT), potential evapotranspiration rate (PER), warmth index (WI), coldness index (CI) and humidity index (HI). The Maxent model was used to predict the potential geographical distribution of the four tree species under current and future scenarios (2041−2060, 2061−2080) SSP126, SSP245 and SSP585. The Jackknife method was used to analyze the main environmental factors affecting its distribution, and the predicted results were tested by the area under receiver operating characteristic curve (AUC).
        Result  (1) Maxent model can well simulate the potential geographical distribution range of major established species in the Loess Plateau, and the average AUC of each species was greater than 0.8. (2) For Pinus sylvestris, Pinus tabuliformis and Caragana korshinskii, both temperature and precipitation limited their distribution, while for Larix gmelinii var. principis-rupprechtii, precipitation was the main factor affecting their distribution. Temperature seasonality, potential evapotranspiration rate, precipitation of the driest month and precipitation of the wettest quarter were the main environmental factors affecting the distribution of Pinus sylvestris. Temperature annual range, temperature seasonality, minimum temperature of the coldest month, precipitation of the warmest quarter and potential evapotranspiration rate were the main factors affecting the distribution of Pinus tabuliformis. The main factors affecting the distribution of Caragana korshinskii are maximum temperature of the warmest month, warmth index, isotherm, potential evapotranspiration rate and precipitation of the coldest season. The main factors affecting the distribution of Larix gmelinii var. principis-rupprechtii were mainly related to precipitation, which were precipitation of the wettest month, precipitation of the wettest quarter, variation coefficient of precipitation, and potential evapotranspiration rate. (3) The potential suitable areas of Pinus tabuliformis, Caragana korshinskii and Larix gmelinii var. principis-rupprechtii will migrate to the northwest, while that of Pinus sylvestris will migrate to the southwest. The potential suitable areas of Pinus tabuliformis and Larix gmelinii var. principis-rupprechtii showed a trend of expanding first and then decreasing, while the potential suitable areas of Caragana korshinskii and Pinus sylvestris would continue to expand, especially the proportion of highly suitable areas of Pinus sylvestris would expand to 50.97% in 2070s.
        Conclusion  Climate change will deprive Pinus tabuliformis and Larix gmelinii var. principis-rupprechtii of some of their highly suitable areas, but at the same time, the highly suitable areas of Caragana korshinskii and Pinus sylvestris will expand significantly. Caragana korshinskii and Pinus sylvestris are preferred in the project of converting farmland to forest land on the Loess Plateau.

       

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