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    日本落叶松人工林叶量动态变化规律及其与数量成熟龄的关系

    Dynamic changing rule in leaf mass of Larix kaempferi plantations and its relationship with quantitative maturity age

    • 摘要:
      目的 探索日本落叶松叶量(叶片干质量)、林分蓄积、林分密度的变化规律,并深入分析林分叶量与数量成熟龄的关系。
      方法 以湖北省建始县长岭岗林场日本落叶松人工林为研究对象,基于30个样地的90株解析木数据,构建了单木叶量、林分密度、林分叶量和林分蓄积模型,探究单木和林分叶量动态变化规律,以及林分叶量高峰时间与数量成熟龄之间的关系。
      结果 (1)添加哑变量(优势木、平均木、劣势木)并以幂函数作为异方差的Logistic方程为单木叶量的最佳模型(R2 = 0.620、rRMSE = 0.554、RMSE = 2.821 kg、MAE = 1.756 kg),林分密度最优模型为Logical方程(R2 = 0.708、rRMSE = 0.190、RMSE = 187.975 株/hm2、MAE = 168.045 株/hm2),林分蓄积最优模型为Korf方程(R2 = 0.703、rRMSE = 0.337、RMSE = 59.681 m3/hm2、MAE = 48.705 m3/hm2)。(2)单木叶量随林龄增加而增加,而林分叶量呈单峰趋势,在24 a达到峰值后下降;不同林层的叶量随林龄增长呈现明显的分层特征。(3)林分叶量与蓄积年平均生长量变化趋势一致,林分叶量达到峰值时间与数量成熟龄(24 a)高度吻合。
      结论 本研究构建的考虑树木等级的叶量预测模型,揭示了单木与林分尺度上的叶量动态差异,研究还发现林分叶量峰值出现时间与数量成熟龄高度一致,为湖北地区日本落叶松的叶量准确预估、科学制定森林经营方案及确定最佳采伐期提供理论依据和实践指导。

       

      Abstract:
      Objective Through growth equations, this paper investigates the dynamics of leaf mass (leaf dry mass), stand volume, and stand density in Larix kaempferi, with a focus on examining the relationship between stand leaf mass and quantitative maturity age.
      Method This study focused on Larix kaempferi plantations in the Changlinggang Forest Farm, Jianshi County, Hubei Province of central China. Based on data from 90 sample trees across 30 sample plots, we developed individual tree leaf mass stand density, stand leaf mass, and stand volume models. The aim was to investigate the dynamic patterns of leaf mass at both individual tree and stand levels, as well as the relationship between peak stand leaf mass timing and quantitative maturity age.
      Result (1) The optimal model for individual tree leaf mass was a heteroscedasticity-adjusted Logistic equation incorporating dummy variables (dominant, intermediate, and suppressed trees) with a power function (R2 = 0.620, rRMSE = 0.554, RMSE = 2.821 kg, MAE = 1.756 kg). The optimal model for stand density was a Logistic equation ( R2 = 0.708, rRMSE = 0.190, RMSE = 187.975 plant/ha, MAE = 168.045 plant/ha). The optimal model for stand volume was a Korf equation (R2 = 0.703, rRMSE = 0.337, RMSE = 59.681 m3/ha, MAE = 48.705 m3/ha). (2) The leaf mass of a single tree increased with forest age, while the leaf mass of stand showed a unimodal trend, reaching its peak at 24 years and then decreasing. The leaf mass of different forest layers showed obvious stratification characteristics with the increase of forest age. (3) The trend of changes in annual average growth of forest leaves and stock volume were consistent, and the time when the forest leaves reaching their peak was highly consistent with the quantity maturity age (24 years).
      Conclusion The tree-class-based leaf mass prediction model developed in this study accurately captures the differences in leaf dynamics at both individual tree and stand scales. It is also found that the peak stand leaf mass coincides closely with quantitative maturity age. These findings provide a theoretical basis and practical guidance for accurate leaf mass estimation, scientific forest management planning, and optimal rotation age determination for Larix kaempferi plantations in Hubei Province of central China.

       

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