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地形和竞争对典型阔叶红松林不同生长阶段树木胸径生长的影响

韩大校 金光泽

韩大校, 金光泽. 地形和竞争对典型阔叶红松林不同生长阶段树木胸径生长的影响[J]. 北京林业大学学报, 2017, 39(1): 9-19. doi: 10.13332/j.1000-1522.20160218
引用本文: 韩大校, 金光泽. 地形和竞争对典型阔叶红松林不同生长阶段树木胸径生长的影响[J]. 北京林业大学学报, 2017, 39(1): 9-19. doi: 10.13332/j.1000-1522.20160218
HAN Da-xiao, JIN Guang-ze. Influences of topography and competition on DBH growth in different growth stages in a typical mixed broadleaved-Korean pine forest, northeastern China[J]. Journal of Beijing Forestry University, 2017, 39(1): 9-19. doi: 10.13332/j.1000-1522.20160218
Citation: HAN Da-xiao, JIN Guang-ze. Influences of topography and competition on DBH growth in different growth stages in a typical mixed broadleaved-Korean pine forest, northeastern China[J]. Journal of Beijing Forestry University, 2017, 39(1): 9-19. doi: 10.13332/j.1000-1522.20160218

地形和竞争对典型阔叶红松林不同生长阶段树木胸径生长的影响

doi: 10.13332/j.1000-1522.20160218
基金项目: 

长江学者和创新团队发展计划项目 IRT_15R09

国家自然科学基金项目 31270473

详细信息
    作者简介:

    韩大校。主要研究方向:森林生态学。Email:550447475@qq.com  地址:150040  黑龙江省哈尔滨市香坊区和兴路51号东北林业大学科技大厦

    责任作者:

    金光泽,博士,教授。主要研究方向:森林生态学。Email:taxus@126.com  地址:同上

  • 中图分类号: S718.5

Influences of topography and competition on DBH growth in different growth stages in a typical mixed broadleaved-Korean pine forest, northeastern China

  • 摘要: 近年来,地形和竞争对树木生长影响机制的研究备受关注。本文基于黑龙江凉水国家级自然保护区内9 hm2典型阔叶红松林动态监测样地,使用2010年和2015年的野外调查数据,分析了地形(坡向、坡度、坡位)和树木间竞争对阔叶红松林主要组成树种胸径生长的影响。结果表明:紫椴、色木槭、冷杉径级Ⅰ(1 cm≤DBH<10 cm)以及红松、紫椴、色木槭、水曲柳径级Ⅱ(DBH≥10 cm)的胸径年均生长量与地形显著相关(P<0.05);竞争对红松、紫椴、色木槭、水曲柳、冷杉、枫桦、青楷槭径级Ⅰ以及红松、紫椴、色木槭、冷杉、春榆径级Ⅱ的胸径年均生长量有显著影响(P<0.05)。总体上,阔叶红松林中耐荫和非耐荫树种组的胸径年均生长量均受地形和竞争的显著影响;阔叶红松林中大径级树木的胸径生长主要受地形影响,而小径级树木的胸径生长主要受竞争影响。

     

  • 图  1  阔叶红松林主要树种胸径与胸径年均生长量关系

    Figure  1.  Relationships between DBH of major tree species and annual average DBH increment in a mixed broadleaved-Korean pine forest

    图  2  地形对阔叶红松林不同生活习性树木胸径年均生长量的影响

    不同大写字母表示不同地形对径级Ⅰ树木胸径年均生长量的影响差异显著(P<0.05),不同小写字母表示不同地形对径级Ⅱ树木胸径年均生长量的影响差异显著(P<0.05)。图 3同此。

    Figure  2.  Influence of topography on annual average DBH increment of different living habits of trees in a mixed broadleaved-Korean pine forest

    Different majuscules represent significant differences of diameter classⅠ's annual average DBH increment at P < 0.05 level on different topography. Different minuscules represent the significant difference of diameter class Ⅱ's annual average DBH increment at P < 0.05 level on different topography. Same as Fig. 3.

    图  3  地形对阔叶红松林不同径级树木胸径年均生长量的影响

    Figure  3.  Influence of topography on annual average DBH increment of different diameter classes of trees in a mixed broadleaved-Korean pine forest

    图  4  竞争压力对耐荫树种组和非耐荫树种组胸径年均生长量的影响

    不同大写字母表示不同竞争压力对耐荫树种组和非耐荫树种组胸径年均生长量的影响差异显著(P<0.05),不同小写字母表示各竞争指数等级中耐荫树种组和非耐荫树种组胸径年均生长量的影响差异显著(P<0.05)。

    Figure  4.  Influence of competition on annual average DBH increment of shade-tolerant and non shade-tolerant groups in a mixed broadleaved-Korean pine forest

    Different majuscules represent the significant difference of annual average DBH increment of shade-tolerant and non-shade groups at P < 0.05 level under different competitive pressures. Different minuscules represent the significant difference of annual average DBH increment of different shade-tolerant groups at P < 0.05 level under same competitive pressures.

    图  5  竞争压力对阔叶红松林不同径级树木胸径年均生长量的影响

    不同大写字母表示竞争压力对径级Ⅰ树木胸径年均生长量的影响差异显著(P<0.05),不同小写字母表示竞争压力对径级Ⅱ树木胸径年均生长量的影响差异显著(P<0.05)。

    Figure  5.  Influence of competition on annual average DBH increment of different diameter classes of trees in a mixed broadleaved-Korean pine forest

    Different majuscules represent the significant difference of annual average DBH increment of diameter classⅠat P < 0.05 level under competitive pressures. Different minuscules represent the significant difference of annual average DBH increment of diameter classeⅡat P < 0.05 level under competitive pressures.

    表  1  地形对阔叶红松林主要组成树种年均胸径生长量的影响

    Table  1.   Influence of topography on annual average DBH increment of major tree species in a mixed broadleaved-Korean pine forest

    mm·a-1
    mm·year-1
    地形
    Topography
    红松Pinus koraiensis紫椴Tilia amurensis色木槭Acer mono水曲柳Fraxinus mandshurica
    坡向
    Slope aspect
    平地Flat1.10±0.182.51±0.18a0.85±0.22b5.38±1.77a0.73±0.07b2.86±0.16a1.27±0.362.76±0.42b
    阴坡Shady slope2.01±0.08b0.80±0.20b0.85±0.27b2.22±0.09b5.80±2.53a
    半阴坡Semi-shady slope0.73±0.151.76±0.12b0.88±0.16b2.09±0.49b0.81±0.10b2.19±0.18b1.60±1.081.94±0.31b
    半阳坡Semi-sunny slope0.75±0.241.90±0.10b1.03±0.12b2.09±0.28b0.89±0.11b2.19±0.07b0.94±0.242.69±0.33b
    阳坡Sunny slope0.52±0.152.04±0.10b2.61±0.16a2.38±0.32b1.33±0.07a2.26±0.07b1.40±0.222.34±0.26b
    坡度
    Slope gradient
    <6°1.04±0.182.38±0.15a0.94±0.18b4.73±0.18a0.85±0.11b2.74±0.13a1.18±0.272.66±0.34
    6°~15°0.75±0.191.92±0.09b1.00±0.12b2.24±0.23b0.81±0.05b2.19±0.05b1.57±0.282.50±0.25
    15°~25°0.70±0.201.99±0.11b0.94±0.16b2.28±0.40b0.89±0.10b2.32±0.15b0.50±0.112.49±0.34
    >25°0.25±0.151.72±0.18b2.64±1.65a2.73±0.34b1.25±0.16a2.12±0.16b1.87±0.622.13±0.52
    坡位
    Slopeposition
    山谷Valley1.04±0.182.45±0.18a1.12±0.27b2.01±1.47a0.75±0.07b2.85±0.15a1.22±0.292.69±0.35
    下坡Lower slope0.71±0.151.92±0.09b1.06±0.10b2.16±0.25b0.73±0.05b2.23±0.07b1.19±0.212.38±0.21
    上坡Upper slope0.64±0.232.07±0.11b0.87±0.11b2.22±0.35b0.75±0.06b2.15±0.07b1.68±0.532.80±0.46
    山脊Ridge1.80±0.04b2.47±0.10a2.10±0.30b1.37±0.59a2.11±0.06b1.40±1.11
    地形
    Topography
    冷杉Abies nephrolepis枫桦Betula costata春榆Ulmus japonica青楷槭Acer tegmentosum
    坡向
    Slope aspect
    平地Flat1.45±0.272.57±0.260.61±0.122.74±0.421.36±0.262.76±1.131.91±0.242.89±0.80
    阴坡Shady slope1.25±0.472.32±0.440.57±0.071.70±0.901.24±0.39
    半阴坡Semi-shady slope1.06±0.202.20±0.230.31±0.072.46±0.241.40±0.204.00±2.202.20±0.191.45±0.26
    半阳坡Semi-sunny slope0.80±0.122.10±0.140.57±0.102.94±0.371.42±0.253.67±0.951.83±0.122.17±0.37
    阳坡Sunny slope1.16±0.132.31±0.160.55±0.082.87±0.291.00±0.123.84±0.881.92±0.212.32±0.39
    坡度Slope gradient<6°1.45±0.222.67±0.280.62±0.112.93±0.331.43±0.233.43±0.741.92±0.182.92±0.66
    6°~15°1.01±0.112.20±0.110.46±0.072.70±0.250.94±0.153.12±0.641.86±0.112.36±0.34
    15°~25°0.82±0.132.13±0.170.56±0.102.68±0.331.21±0.203.33±0.722.12±0.201.64±0.20
    >25°1.14±0.192.23±0.360.20±0.092.55±0.561.28±0.291.92±0.531.74±0.97
    坡位Slope position山谷Valley1.58±0.25a2.54±0.250.60±0.122.99±0.411.31±0.224.03±1.571.94±0.222.89±0.80
    下坡Lower slope0.96±0.10b2.32±0.120.51±0.072.76±0.221.22±0.143.80±0.601.91±0.102.07±0.28
    上坡Upper slope1.03±0.13b1.98±0.140.48±0.082.68±0.330.93±0.272.44±1.222.01±0.232.16±0.41
    山脊Ridge2.72±1.060.48±0.082.55±0.560.87±0.77
    注:Ⅰ表示1 cm≤DBH<10 cm,Ⅱ表示DBH≥10 cm;表中数值为平均值±标准误;不同小写字母表示不同地形下差异显著(P<0.05)。Notes:Ⅰ, 1 cm≤DBH<10 cm; Ⅱ, DBH≥10 cm; Data in table is mean±SE; Different minuscules represent significant difference at P<0.05 level on different topography conditions.
    下载: 导出CSV

    表  2  竞争对阔叶红松林主要组成树种年均胸径生长量的影响

    Table  2.   Influence of competition on annual average DBH increment of major tree species in a mixed broadleaved-Korean pine forest

    mm·a-1
    mm·year-1
    竞争指数Competition index(CI)红松Pinus koraiensis紫椴Tilia amurensis色木槭Acer mono水曲柳Fraxinus mandshurica
    <52.34±0.06a2.12±0.07a1.87±0.13a2.91±0.39a1.36±0.06a2.46±0.13a2.38±0.69a2.56±0.18
    5~100.88±0.14b1.47±0.13b1.09±0.13b1.69±0.26b1.11±0.08b1.44±0.15b1.39±0.22b1.40±0.55
    >100.98±0.15b1.71±0.25b0.99±0.08b1.60±0.10b0.81±0.43c1.11±0.42b0.86±0.18b
    竞争指数Competition index(CI)冷杉Abies nephrolepis枫桦Betula costata春榆Ulmus japonica青楷槭Acer tegmentosum
    <52.03±0.07a2.38±0.10a1.92±0.16a2.80±0.171.25±0.342.59±0.37a3.09±0.42a2.38±0.26
    5~101.11±0.08b1.50±0.16b1.05±0.10b2.13±0.711.28±0.161.31±0.14b2.24±0.25b2.16±0.22
    >101.01±0.11b1.43±0.39b0.60±0.08c1.23±0.181.08±0.12b1.79±0.09c2.08±0.20
    注:Ⅰ表示1 cm≤DBH<10 cm,Ⅱ表示DBH≥10 cm;不同小写字母表示不同竞争指数等级下差异显著(P<0.05)。Notes:Ⅰ, 1 cm≤DBH<10 cm; Ⅱ, DBH≥10 cm; Different minuscules represent significant difference at P<0.05 level under different competition index grades.
    下载: 导出CSV
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  • 收稿日期:  2016-06-30
  • 修回日期:  2016-10-29
  • 刊出日期:  2017-01-01

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