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银杏生物量分配格局及异速生长模型

刘坤 曹林 汪贵斌 曹福亮

刘坤, 曹林, 汪贵斌, 曹福亮. 银杏生物量分配格局及异速生长模型[J]. 北京林业大学学报, 2017, 39(4): 12-20. doi: 10.13332/j.1000-1522.20160374
引用本文: 刘坤, 曹林, 汪贵斌, 曹福亮. 银杏生物量分配格局及异速生长模型[J]. 北京林业大学学报, 2017, 39(4): 12-20. doi: 10.13332/j.1000-1522.20160374
LIU Kun, CAO Lin, WANG Gui-bin, CAO Fu-liang. Biomass allocation patterns and allometric models of Ginkgo biloba[J]. Journal of Beijing Forestry University, 2017, 39(4): 12-20. doi: 10.13332/j.1000-1522.20160374
Citation: LIU Kun, CAO Lin, WANG Gui-bin, CAO Fu-liang. Biomass allocation patterns and allometric models of Ginkgo biloba[J]. Journal of Beijing Forestry University, 2017, 39(4): 12-20. doi: 10.13332/j.1000-1522.20160374

银杏生物量分配格局及异速生长模型

doi: 10.13332/j.1000-1522.20160374
基金项目: 

林业公益性行业科研专项 201504105

详细信息
    作者简介:

    刘坤,博士生。主要研究方向:森林培育。Email:vaguelk@outlook.com  地址:210037 江苏省南京市龙蟠路159号南京林业大学林学院

    责任作者:

    曹林,讲师。主要研究方向:林业遥感。Email: ginkgocao@gmail.com  地址:同上

  • 中图分类号: S792.95

Biomass allocation patterns and allometric models of Ginkgo biloba

  • 摘要: 以苏北地区银杏人工林为研究对象,选取13株进行整株挖掘,分析不同器官生物量的分配格局,以及地上和地下生物量之间的关系;再分别以胸径(D)、树高(H)、D2HDaHb为自变量建立银杏各器官生物量模型,选择调整决定系数(Radj2)、残差平方和(SSE)、平均偏差(ME)、平均绝对偏差(MAE)和平均相对误差(MPE)作为选择最优模型的检验指标,根据检验结果筛选出各器官的最优模型。结果表明:13株银杏的整株生物量变化范围为28.50~320.27 kg,树干生物量占总生物量的49.4%~56.6%,树枝生物量占总生物量的12.1%~18.9%,树叶生物量占总生物量的3.8%~5.5%,根生物量占总生物量的26%;地上部分生物量与地下生物量线性方程的斜率为0.35,具有显著的线性相关性(P<0.01);枝和叶生物量都集中于树冠中部,树冠上层和下层的枝、叶生物量明显低于树冠中层生物量(P<0.05),上层和下层生物量之间差异不显著(P>0.05),70%根生物量集中0~1.0 m的土层;枝水平上,基于基径和枝长的枝生物量模型解释量超过95%;在各器官生物量最优模型选择上,以D为自变量的W=aDb的叶、枝、地上部分生物量模型要优于其他模型;树干、根和全株生物量则是以W=aDbHc模型最优。银杏各器官生物量表现为干>根>枝>叶,枝和叶生物量垂直分配上,中冠层占最大比例;基于树高和胸径的相对生长模型可以实现对银杏各器官生物量的准确拟合,银杏生物量及碳储量的有效估算。

     

  • 图  1  银杏地上生物量与地下生物量之间的关系

    Figure  1.  Relationship between aboveground and belowground biomass of Ginkgo biloba

    图  2  银杏树干、树枝、树叶、树根生物量空间分布图

    Figure  2.  Horizontal and vertical distribution of stem, branch, leaf and root biomass of G. biloba

    表  1  标准木基本信息表

    Table  1.   Basic characteristics of sample trees

    径级
    Diameter at breast height (DBH) class/cm
    株数
    Tree number
    胸径
    DBH/cm
    树高
    Tree height(H)/m
    树冠长度
    Grown length/m
    南北向冠幅
    South-north crown width/m
    东西向冠幅
    East-west crown width/m
    生物量
    Biomass/kg
    10~15710.911.16.62.63.328.50
    11.411.29.33.22.335.70
    13.312.310.25.36.456.05
    13.412.09.66.05.056.61
    13.511.47.84.94.852.35
    13.611.28.14.94.559.46
    13.811.79.34.74.360.26
    15~20317.113.911.93.98.881.36
    17.713.59.65.15.6102.84
    18.812.610.25.76.3108.95
    >20320.113.011.56.07.2134.82
    24.814.512.06.58.6223.20
    27.213.612.68.49.7320.27
    下载: 导出CSV

    表  2  株水平上各器官生物量测定值

    Table  2.   Measurement of each component biomass at the tree level

    kg
    组分
    Component
    径级DBH class
    10~15 cm15~20 cm>20 cm
    叶生物量
    Leaf biomass
    2.623.8017.96
    干生物量
    Stem biomass
    28.2051.59146.00
    枝生物量
    Branch biomass
    6.0216.0172.60
    地上部分生物量
    Aboveground biomass
    36.8471.40236.56
    根生物量
    Root biomass
    13.0126.3283.70
    总生物量
    Total biomass
    49.8597.72320.26
    下载: 导出CSV

    表  3  枝水平上枝生物量异速生长模型

    Table  3.   Allometric models for branch at branch level

    生物量
    Biomass/kg
    模型
    Model
    系数CoefficientRadj2显著性
    Sig.
    CFSSEMAEMPE
    abc
    枝BranchlnW=a+blnBD-0.46 ns3.039***-0.904P<0.0011.0781.6990.27924.718
    lnW=a+blnBD+ClnBL0.039 ns0.916 ns2.1290.957P<0.0011.0260.7150.20112.038
    注:*表示在0.05水平上差异显著;**表示在0.01水平上差异显著;***表示在0.001水平上差异显著;ns表示在0.05水平上差异不显著;BD表示基径;BL表示枝长。下同。Notes: * means significant difference at P<0.05 level; ** means significant difference at P<0.01 level; *** means significant difference at P<0.01 level; ns means no significant difference at P<0.05 level; BD means branch diameter; BL means branch length. The same below.
    下载: 导出CSV

    表  4  银杏不同器官生物量模型参数估计、拟合结果和检验结果

    Table  4.   Parameter estimation, fitting and test results of different components of G. biloba biomass

    模型
    Model
    生物量
    Biomass/kg
    系数CoefficientRadj2显著性Sig.CFSSEMAEMPE
    abc
    lnW=a+blnD叶Leaf-5.21***2.37***0.868P<0.0011.025 00.7520.19917.535
    干Stem-2.23***2.17***0.989P<0.0011.000 10.0460.0481.280
    枝Branch-6.95***3.36***0.871P<0.0011.077 01.4710.26326.879
    地上Aboveground-2.56***2.40***0.980P<0.0011.003 00.1040.0731.848
    根Root-3.75***2.45***0.989P<0.0011.000 60.1070.0782.761
    全株Total-2.29***2.41***0.985P<0.0011.000 20.0810.0661.550
    lnW=a+blnH叶Leaf-14.09**6.13**0.571P<0.0101.075 42.5180.35633.011
    干Stem-10.75**5.77***0.698P<0.0011.004 71.6340.2977.758
    枝Branch-19.13**8.53**0.550P<0.0011.153 16.1830.48944.667
    地上Aboveground-10.51**5.09***0.668P<0.0011.005 42.1660.3257.890
    根Root-13.24**6.46***0.678P<0.0011.010 82.2650.33711.212
    全株Total-11.55**6.32***0.068P<0.0011.004 82.1590.3277.393
    lnW=a+bln(D2H)叶Leaf-6.95***1.03***0.853P<0.0011.025 40.7700.20617.974
    干Stem-3.84***0.95***0.980P<0.0011.000 20.0800.0581.556
    枝Branch-9.38***1.46***0.852P<0.0011.080 01.7190.28828.092
    地上Aboveground-4.32***1.04***0.967P<0.0011.000 50.1680.0822.040
    根Root-5.60***1.07***0.967P<0.0011.001 00.1740.1033.542
    全株Total-4.07***1.05***0.972P<0.0011.000 30.1450.0801.823
    lnW=a+blnD+clnH叶Leaf-3.26***2.68***-1.12***0.861P<0.0011.025 20.7490.20017.649
    干Stem-1.61***2.27***-0.36***0.990P<0.0011.000 20.0460.4801.277
    枝Branch-3.14***3.97***-2.18***0.870P<0.0011.077 21.3770.25726.786
    地上Aboveground-1.18***2.62***-0.79***0.981P<0.0011.000 30.1000.0731.864
    根Root-2.64***2.63***-0.64***0.980P<0.0011.000 70.1040.0742.657
    全株Total-1.01***2.61***-0.73***0.986P<0.0011.000 20.0770.0671.564
    下载: 导出CSV
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  • 收稿日期:  2016-11-16
  • 修回日期:  2017-02-21
  • 刊出日期:  2017-04-01

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