Species diversity of Larix principis-rupprechtii communities in Shanxi Province of northern China
-
摘要: 本文以华北落叶松群落为研究对象,在山西境内共调查了40个典型样地,应用数量生态学的相关计算方法对华北落叶松群落进行了TWINSPAN分类,依据不同的指示种将40个典型样地划分成为10个群丛。通过运用Patrick物种丰富度指数(R)、Simpson指数(D)、Shannon-Wiener指数(H′)和Pielou均匀度指数(E)分别研究了山西华北落叶松群落的物种多样性。由于群落H′指数受到R指数和E指数共同影响,因此H′指数含有较多群落组成结构、分布状况的信息,H′指数的结果表明:10个群丛物种的多样性指数顺序为群丛2>群丛1>群丛3>群丛9>群丛4>群丛5>群丛10>群丛6>群丛7>群丛8。10个群丛的物种多样性指数均较高,且存在较大差异,表明了山西华北落叶松群落的结构复杂,组织水平高,生长环境适宜。3个层片的多样性指数总体上呈现出草本层>灌木层>乔木层,说明草本层的物种最为丰富, 对于群落总体多样性的贡献最为突出。各多样性指数的方差分析结果表明, 10个群丛的指数R差异显著(P < 0.05),H′差异极显著(P < 0.01),而D和E两者的差异性并不显著(P>0.05)。Abstract: In order to explore the species diversity of Larix principis-rupprechtii forests in Shanxi Province of northern China, 40 typical sample plots of Larix principis-rupprechtii forests were investigated from the field. Based on the result of TWINSPAN classification, the communities were classified into 10 associations. The species diversity of Larix principis-rupprechtii communities in Shanxi was studied by species richness index (R), Simpson index (D), Shannon-Wiener index (H′) and Pielou evenness index (E). Through this study, it is hopeful to provide scientific basis for protection and rational exploitation of Larix principis-rupprechtii communities. Community H′ index was affected by R and E indexes together, containing more information about community composition structure and distribution condition. The results indicated that the rank of species richness index for ten associations was association 2>association 1>association 3>association 9>association 4>association 5>association 10>association 6>association 7>association 8. All Larix principis-rupprechtii associations had higher species diversity index, and it had big difference in the different associations. It showed that the Larix principis-rupprechtii communities had more complex structure, higher organization level and appropriate growth environment. The biodiversity index was the highest in herb layer, followed by the shrub layer and the forest layer, which suggested that the herb layer contributed to the overall community richness mostly. The results of one-way ANOVA for those indices among 10 associations showed that R had a significant difference (P < 0.05), and H′ had extremely significant difference (P < 0.01), while the difference between D and E was not significant (P>0.05).
-
Keywords:
- Shanxi Province /
- Larix principis-rupprechtii /
- species diversity
-
-
表 1 山西华北落叶松群落物种多样性指数方差分析结果
Table 1 ANOVA for the species diversity indices of Larix principis-rupprechtii communities in Shanxi Province
多样性指数
Diversity index组间平方和
Square sum among classes (SSa)组间dfdf
amongclasses组内平方
和Square sum within class (SSe)组内dfdf
withinclass组间均方和
Mean square sum among classes (MSa)组内均方和
Mean square sum within class (MSe)F P R 1 011.860 7 1 407.851 30 144.551 46.928 3.080 0.014 D 0.091 7 0.174 30 0.013 0.006 2.257 0.057 H′ 2.288 7 1.945 30 0.327 0.065 5.043 0.001 E 0.069 7 0.267 30 0.010 0.009 1.111 0.382 -
[1] 李文荣, 齐力旺.山西华北落叶松天然林的地理分布和种群变异规律的研究[J].林业科学, 1992, 6(12):493-501. http://www.cnki.com.cn/Article/CJFDTotal-LYKE199206001.htm LI W R, QI L W. A study on geographic distribution and population variation law of Larix principis-rupprechtii Mayr natural forest in Shanxi[J]. Scientia Silvae Sinicea, 1992, 6(12):493-501. http://www.cnki.com.cn/Article/CJFDTotal-LYKE199206001.htm
[2] 赵匡记, 王利东, 王立军, 等.华北落叶松蓄积量及生产力研究[J].北京林业大学学报, 2015, 37(2):24-31. doi: 10.13332/j.cnki.jbfu.2015.02.011 ZHAO K J, WANG L D, WANG L J, et al. Stock volume and productivity of Larix principis-rupprechtii in northern and northwestern China[J]. Journal of Beijing Forestry University, 2015, 37(2):24-31. doi: 10.13332/j.cnki.jbfu.2015.02.011
[3] TOMMASO J, MARTA C. Going beyond taxonomic diversity: deconstructing biodiversity pattern sreveals the true cost of ice plant invasion[J]. Diversity and Distributions, 2013, 19:1566-1577. doi: 10.1111/ddi.12124
[4] 黄忠良, 孔国辉, 何道泉.鼎湖山植物群落多样性的研究[J].生态学报, 2000, 20(2):193-198. http://d.old.wanfangdata.com.cn/Periodical/stxb200002004 HUANG Z L, KONG G H, HE D Q. Plant community diversity in Dinghushan Nature Reserve[J]. Acta Ecologica Sinica, 2000, 20(2):193-198. http://d.old.wanfangdata.com.cn/Periodical/stxb200002004
[5] 马克平, 黄建辉, 于顺利, 等.北京东灵山地区植物群落多样性的研究[J].生态学报, 1995, 15(3): 268-277. doi: 10.3321/j.issn:1000-0933.1995.03.006 MA K P, HUANG J H, YU S L, et al. Plant community diversity in Donglingshan Mountain, Beijing, China[J] Acta Ecologica Sinica, 1995, 15(3): 268-277. doi: 10.3321/j.issn:1000-0933.1995.03.006
[6] HARTNETT D C, WILSON W T. Mycorrhizae influence plant community structure and diversity in tallgrass prairie[J]. Ecology, 1999, 80: 1187-1195. doi: 10.1890/0012-9658(1999)080[1187:MIPCSA]2.0.CO;2
[7] 李旭华.晋南地区植被与环境关系分析[D].太原: 山西大学, 2014. LI X H. Analysis on the relationship between vegetation and environment in Southern Shanxi[D].Taiyuan: Shanxi University, 2014.
[8] 方精云, 王襄平, 沈泽昊, 等.植物群落清查的主要内容、方法和技术规范[J].生物多样性, 2009, 17(6): 533-548. http://www.wanfangdata.com.cn/details/detail.do?_type=perio&id=swdyx200906002 FANG J Y, WANG X P, SHEN Z H, et al. Methods and protocols for plant community inventory[J]. Biodiversity Science, 2009, 17(6):533-548. http://www.wanfangdata.com.cn/details/detail.do?_type=perio&id=swdyx200906002
[9] 赵小娜, 秦晓娟, 董刚, 等.庞泉沟自然保护区植物群落分类学多样性[J].应用生态学报, 2014, 25(12):3437-3442. http://d.old.wanfangdata.com.cn/Periodical/yystxb201412008 ZHAO X N, QIN X J, DONG G, et al. Taxonomic diversity of the plant communities in Pangquangou Nature Reserve, Shanxi Province, China[J]. Chinese Journal of Applied Ecology, 2014, 25(12):3437-3442. http://d.old.wanfangdata.com.cn/Periodical/yystxb201412008
[10] 张金屯.数量生态学[M].北京:科学出版社, 2004. ZHANG J T. Quantitative ecology[M].Beijing: Science Press, 2004.
[11] 马克平.生物多样性的测度方法:α多样性的测度方法(上)[J].生物多样性, 1994, 2(3): 162-168. doi: 10.3321/j.issn:1005-0094.1994.03.007 MA K P. Biological diversity measure: α diversity measure method[J]. Biodiversity Science, 1994, 2(3):162-168. doi: 10.3321/j.issn:1005-0094.1994.03.007
[12] 吴征镒.中国植被[M].北京:科学出版社, 1995. WU Z Y. Chinese vegetation[M].Beijing: Science Press, 1995.
[13] 张峰, 张金屯, 上官铁梁.历山自然保护区猪尾沟森林群落植物多样性研究[J].植物生态学报, 2002, 26(增刊): 46-51. http://d.old.wanfangdata.com.cn/Periodical/zwstxb2002z1006 ZHANG F, ZHANG J T, SHANGGUAN T L. Plant diversity of forest community in Zhuweigou of Lishan Mountain Nature Reserve[J]. Chinese Journal of Plant Ecology, 2002, 26(Suppl.): 46-51. http://d.old.wanfangdata.com.cn/Periodical/zwstxb2002z1006
[14] 李旭华, 邓永利, 张峰, 等.山西庞泉沟自然保护区森林群落物种多样性[J].生态学杂志, 2013, 32(7):1667-1673. http://d.old.wanfangdata.com.cn/Periodical/stxzz201307003 LI X H, DENG Y L, ZHANG F, et al. Species diversity of forest communities in Pangquangou Nature Reserve, Shanxi of China[J]. Chinese Journal of Ecology, 2013, 32(7):1667-1673. http://d.old.wanfangdata.com.cn/Periodical/stxzz201307003
[15] 郭华, 张桂萍, 铁军, 等.太行山南段油松群落物种多样性研究[J].植物科学学报, 2015, 32(2):151-157. http://d.old.wanfangdata.com.cn/Periodical/whzwxyj201502004 GUO H, ZHANG G P, TIE J, et al. Analysis of species diversity of Pinus tabulaeformis forest community in southern Taihang Mountains[J]. Plant Science Journal, 2015, 32(2):151-157. http://d.old.wanfangdata.com.cn/Periodical/whzwxyj201502004
[16] 张丽, 廉凯敏, 赵璐璐, 等.太宽河自然保护区栓皮栎群落物种多样性研究[J].山西大学学报(自然科学版), 2010, 33(2):296-301. http://d.old.wanfangdata.com.cn/Periodical/sxdxxb201002031 ZHANG L, LIAN K M, ZHAO L L, et al. Species diversity of Quercus variabilis communities in Taikuanhe Nature Reserve, Shanxi[J], Journal of Shanxi University (Natural Science Edition), 2010, 33(2):296-301. http://d.old.wanfangdata.com.cn/Periodical/sxdxxb201002031
[17] 石慧, 王孝安, 郭华.秦岭华北落叶松人工林群落结构及物种多样性[J].安徽农学通报, 2008, 14(15):159-204. doi: 10.3969/j.issn.1007-7731.2008.15.074 SHI H, WANG X A, GUO H. Community structure and species diversity of Larix principis-rupprechtii plantation in Qinling Mountains[J]. Anhui Agricultural Science Bulletin, 2008, 14(15):159-204. doi: 10.3969/j.issn.1007-7731.2008.15.074
[18] 陈东莉, 郭晋平, 杜宁宁.间伐强度对华北落叶松林下生物多样性的影响[J].东北林业大学学报, 2011, 39(4):37-38. doi: 10.3969/j.issn.1000-5382.2011.04.012 CHEN D L, GUO J P, DU N N. Effect of thinning intensity on biodiversity of undergrowth vegetation in a Larix principis-rupprechtii plantation[J].Journal of Northeast Forestry University, 2011, 39(4):37-38. doi: 10.3969/j.issn.1000-5382.2011.04.012
[19] 尹文珂, 胡理乐, 卢希, 等.秦岭中段油松天然次生林群落物种多样性研究[J].西北农林科技大学学报(自然科学版), 2016, 33(5):55-63. http://d.old.wanfangdata.com.cn/Periodical/xbnydxxb201605008 YIN W K, HU L L, LU X, et al. Species diversity of Pinus tabuliformis natural secondary forest in Qinling Mountains[J]. Journal of Northwest A & F University(Natural Science Edition), 2016, 33(5):55-63. http://d.old.wanfangdata.com.cn/Periodical/xbnydxxb201605008
[20] 王俊玲, 金红喜, 杨占彪, 等.六盘山华北落叶松人工林多样性、生产力研究[J].兰州大学学报(自然科学版), 2008, 44(1):31-35. doi: 10.3321/j.issn:0455-2059.2008.01.007 WANG J L, JIN H X, YANG Z B, et al. Species diversity and productivity of Larix principis-rupprechtii plantation woods in Liupan Mountains[J].Journal of Lanzhou University(Natural Science Edition), 2008, 44(1):31-35. doi: 10.3321/j.issn:0455-2059.2008.01.007
-
期刊类型引用(6)
1. 林秀云,孙圆,刘晨曦,姚睿涵,周春国,曹林,曹福亮. 依据地面激光扫描数据的杉木材积建模与造材. 东北林业大学学报. 2022(01): 33-39 . 百度学术
2. 李沛婷,赵庆展,田文忠,马永建. 结合无人机载LiDAR点云法向量的K-means++聚类精简. 国土资源遥感. 2020(02): 103-110 . 百度学术
3. 程子阳,任国全,张银. 扫描线段特征用于三维点云地面分割. 光电工程. 2019(07): 111-120 . 百度学术
4. 蔡越,徐文兵,梁丹,邓愫愫,李翀. 基于激光回波强度判别毛竹年龄. 中国激光. 2018(01): 272-280 . 百度学术
5. 曾碧,黄文. 一种融合多特征聚类集成的室内点云分割方法. 计算机工程. 2018(03): 281-286 . 百度学术
6. 田青华,白瑞林,李杜. 基于改进欧氏聚类的散乱工件点云分割. 激光与光电子学进展. 2017(12): 316-324 . 百度学术
其他类型引用(7)