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边材生理机能及心材形成机理的研究进展

罗蓓 何蕊 杨燕

罗蓓, 何蕊, 杨燕. 边材生理机能及心材形成机理的研究进展[J]. 北京林业大学学报, 2018, 40(1): 120-129. doi: 10.13332/j.1000-1522.20170295
引用本文: 罗蓓, 何蕊, 杨燕. 边材生理机能及心材形成机理的研究进展[J]. 北京林业大学学报, 2018, 40(1): 120-129. doi: 10.13332/j.1000-1522.20170295
Luo Bei, He Rui, Yang Yan. A review of physiological function of sapwood and formation mechanism of heartwood[J]. Journal of Beijing Forestry University, 2018, 40(1): 120-129. doi: 10.13332/j.1000-1522.20170295
Citation: Luo Bei, He Rui, Yang Yan. A review of physiological function of sapwood and formation mechanism of heartwood[J]. Journal of Beijing Forestry University, 2018, 40(1): 120-129. doi: 10.13332/j.1000-1522.20170295

边材生理机能及心材形成机理的研究进展

doi: 10.13332/j.1000-1522.20170295
基金项目: 

国家自然科学基金项目 31700481

国家自然科学基金项目 31570555

国家自然科学基金项目 31360157

详细信息
    作者简介:

    罗蓓,博士,讲师。主要研究方向:木材解剖学。Email:49346467@qq.com 地址: 650224 云南省昆明市盘龙区白龙路白龙寺300号西南林业大学200号信箱

  • 中图分类号: S718.47

A review of physiological function of sapwood and formation mechanism of heartwood

  • 摘要: 树木的边材是木质部内具有生理功能的组织,心材虽无生理功能,但对木材的利用却有非常重要的影响。树木心材形成机制是木材科学中几个尚未完全了解的问题之一。本文总结了有关边材生理机能与心材形成机理研究的重点成果,据此来讨论特异性带色心材的形成及调控机制。鉴于含水率对边材薄壁细胞生理机能的重要性,通过揭示木射线组织如何在心边材中间区调控水分导致心材形成的机理,为心材的人工调控提供一定的理论基础。

     

  • 图  1  美东红柏(Juniperus virginiana)边心材中间区的白线带

    Figure  1.  The white line at the boundary between sapwood and heartwood of Juniperus virginiana

    图  2  美东红柏白线带因圆盘干燥而消失

    Figure  2.  Disapperance of the white line in Juniperus virginiana due to disk drying

    图  3  美洲红杉(Sequoia sempervirens)边材射线薄壁细胞显示其内的淀粉粒和细胞核[38]

    Figure  3.  Redwood sapwood ray parenchyma showing starch grains and nuclei in ray cells[38]

    图  4  在美洲红杉边心材中间区薄壁细胞内的淀粉粒被转化成抽提物浸入紧邻管胞内[38]

    Figure  4.  Conversion of starch grains in redwood sapwood ray parenchyma into extractives, and permeation of extractives from ray cells into adjacent tracheids[38]

    图  5  美洲红杉刚采伐时的边/心材界线(箭头所示)及储存期间新增加的心材[36]

    Figure  5.  Sapwood/heartwood boundary (as shown by arrows) of redwood at the time of tree felling and newly formed heartwood during storage[36]

    图  6  针叶树材应压力的偏心生长轮和偏心心材[3]

    Figure  6.  Eccentric annual rings and heartwood in compression wood[3]

    图  7  不同种杨树因高湿木造成的非特性带色心材

    Figure  7.  Facultatively colored heartwood in different species of poplar stimulated by wetwood

    图  8  沉香的主要倍半萜烯类抽提物

    Figure  8.  Major sesquiterpenes in agarwood

    表  1  常见树木边材里生长轮的数目[3]

    Table  1.   Number of growth rings in the sapwood of different wood species[3]

    针叶树材Conifer wood生长轮数Growth
    ring number
    阔叶树材Broadleaved wood生长轮数Growth
    ring number
    柳杉Crytomeria japonica D. Don6~8梓树Catalpa bignoniodes Walt.1~2
    湿地松Pinus elliotti Englm.8~12刺槐Robinia pseudoacacia L.2~3
    北美香柏Thuja occidentalis L.10~15灰胡桃Juglans cinerea L.5~6
    胶冷杉Abies balsamea (L) Mill.10~20山毛榉Fagus grandifolia Ehrh.20~30
    放射松Pinus radiata D. Don12~16颤杨 Populus tremuloides Michx.25~30
    北美落叶松Larix larcina (Du Roi) K. Koch15~25银槭Acer saccharum Marsh.30~40
    班克松 Pinus banksiana Lamb.20~35糖槭Acer saccharinum L.40~50
    北美柱松Pinus contorta Englem.25~50黑桦Betula nigra L.40~50
    欧洲赤松Pinus sylvestris L.25~70枫香Liquidambar styraciflua L.60~70
    庞克松 Pinus ponderosa Laws>100山茱萸Nyssa sylvastica Marsh.80~100
    下载: 导出CSV
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  • 收稿日期:  2017-08-22
  • 修回日期:  2017-11-10
  • 刊出日期:  2018-01-01

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