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    周来, 程小芳, 张梦弢. 林窗对华北落叶松天然次生林林下物种多样性的影响[J]. 北京林业大学学报. DOI: 10.12171/j.1000-1522.20220223
    引用本文: 周来, 程小芳, 张梦弢. 林窗对华北落叶松天然次生林林下物种多样性的影响[J]. 北京林业大学学报. DOI: 10.12171/j.1000-1522.20220223
    Zhou Lai, Cheng Xiaofang, Zhang Mengtao. Effects of forest gaps on understory species diversity of Larix principis-rupprechtii natural secondary forest[J]. Journal of Beijing Forestry University. DOI: 10.12171/j.1000-1522.20220223
    Citation: Zhou Lai, Cheng Xiaofang, Zhang Mengtao. Effects of forest gaps on understory species diversity of Larix principis-rupprechtii natural secondary forest[J]. Journal of Beijing Forestry University. DOI: 10.12171/j.1000-1522.20220223

    林窗对华北落叶松天然次生林林下物种多样性的影响

    Effects of forest gaps on understory species diversity of Larix principis-rupprechtii natural secondary forest

    • 摘要:
      目的 研究华北落叶松天然次生林林窗大小和光因子对林下植被多样性的影响,为森林物种多样性的保护和可持续经营提供科学依据。
      方法 以关帝山华北落叶松天然次生林为研究对象,按面积等级选取12个林窗,于各林窗内设置5 m × 5 m样方3个,以调查样方中的灌木,同时在每个5 m × 5 m样方内设置1 m × 1 m 的草本调查样方1个,再设置12个面积相同的林内对照样方(CK),以同样的方法调查植被。应用Simpson优势度指数(H′)、Shannon-Wiener指数(H)、Pielou均匀度指数(Jsw)、Margalef丰富度指数(D)4种多样性指标,分析华北落叶松天然次生林不同面积等级林窗中物种组成及物种多样性的差异。调查不同面积林窗的光因子指标,分析华北落叶松天然次生林不同面积林窗中光因子与物种多样性的关系。
      结果 在调查的12个林窗中,共记录到灌木植物8种,草本植物32种。林窗草本层的物种多样性指数均大于林内,而灌木层的物种多样性指数仅有Ⅲ级林窗和Ⅳ级林窗大于林内;对于灌木层,随着林窗面积的增大,H值呈现先增高后降低趋势,其中Ⅲ级林窗内的H值最大,Ⅰ级林窗内的H值最小,均大于CK,但草本层的H′指数与林窗面积存在显著相关性(P < 0.05)。对于草本层,H、H′、JswD在小林窗最大,随林窗面积增大而逐渐减小,但均大于CK。草本层的D值较灌木层高,其随林窗面积增加的变化趋势比灌木层更加明显。林窗内的直射辐射量、散射辐射量和总辐射量随着林窗面积的增大而显著变化(P < 0.05)。随着林窗面积的增大散射辐射量和总辐射量也随之增大,而直接辐射量是随着林窗的面积增大先减小后增大,在林窗面积为50 ~ 100 m2时,直接辐射量最小。
      结论 林窗面积在100 ~ 150 m2时有利于提升灌木层的物种多样性水平,在0 ~ 50 m2时有利于提升草本层的物种多样性水平,而在50 ~ 100 m2林下群落物种丰富度最高;林窗内适宜的光因子有利于增加林下植被物种多样性,而过高的光因子反而不利于林下植被多样性的形成。华北落叶松天然次生林林下物种多样性保护过程中,应以中小面积的林窗为主。

       

      Abstract:
      Objective The aim of this study was to study the effects of gap size and light factor of gaps on understory plant diversity in the Larix principis-rupprechtii natural secondary forest.
      Method Taking the natural secondary Larix principis-rupprechtii forest in Guandi Mountain as the research object, 12 gaps were selected according to the area grade, and three 5 m × 5 m quadrats were set in each gap to investigate the shrubs in the quadrats. Each 5 m × 5 m quadrat was set up with a 1 m × 1 m herb investigation quadrat, and another 12 forest control check (CK) with the same area were set up to investigate vegetation in the same way. Four diversity indicators, Simpson dominance index, Shannon-Wiener index, Pielou evenness index, and Margalef richness index, were used to analyze the differences in species composition and species diversity in different area grades of the Larix principis-rupprechtii natural secondary forest. Investigate the light factor indicators of different areas of forest gaps, and analyze the relationship between light factors and species diversity in different areas of Larix principis-rupprechtii natural secondary forest.
      Result In the 12 forest gaps investigated, a total of 8 species of shrubs and 32 species of herbaceous plants were recorded. The species diversity index of the herb layer in the gap is larger than that in the forest, while the species diversity index of the shrub layer is only higher than the forest gap of grade Ⅲ and grade Ⅳ. In the shrub layer, with the increase of the gap area, the H value of the Shannon-Wiener index shows a trend of first increasing and then decreasing. The H value is the smallest, and both are greater than CK, but there was a significant correlation between the H′ index of the herb layer and the gap area (P < 0.05). In the herb layer, the Shannon-Wiener index H, Simpson index H′, Pielou evenness index Jsw, and Margalef abundance index D were the largest in the small gap, and gradually decreased with the increase of the gap area, but it was higher than CK. The D value of the herb layer is higher than that of the shrub layer, and the change trend of the D value of the herb layer with the increase of the gap area is more obvious than that of the shrub layer. The direct radiation, scattered radiation and total radiation in the gap changed significantly with the increase of the gap area (P < 0.05). With the increase of the gap area, the amount of scattered radiation and total radiation also increases, and the amount of direct radiation is reduced first and then increased as the area of the forest gap increases, and the direct radiation is the smallest when the forest gap area is 70 ~ 100 m2.
      Conclusion When the forest gap area is 100 ~ 150 m2, it is beneficial to improve the species diversity level of the shrub layer, and it is beneficial to improve the species diversity level of the herb layer when it is 0 ~ 50 m2, and the species richness of the understory community is the highest in 50 ~ 100 m2; Appropriate light factors in forest gaps are beneficial to increase the diversity of understory vegetation species, however, too high light factor is not conducive to the formation of understory vegetation diversity. Therefore, in the process of protection of forest species diversity of Larix principis-rupprechtii natural secondary forests, small and medium-sized gaps should be the focuses.

       

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